Sussex centre Billy Twelvetrees has retained his place in the England squad for the Autumn internationals next month.

Fellow Sussex players Joe Marler and Joe Launchbury are also in the squad for the Test matches against New Zealand, South Africa, Samoa and Australia.

Twelvetrees, from Pulborough, keeps his place despite enduring a mixed tour to New Zealand in the summer.

High-profile omissions from the 33 are Danny Cipriani and Freddie Burns with Owen Farrell, George Ford and Stephen Myler chosen as the squad's three fly-halves.

England's squad will meet at Pennyhill Park on Sunday ahead of the opening fixture against New Zealand on November 8 and coach Stuart Lancaster confirmed there will be further additions.

''We are still monitoring a number of players injury-wise,'' Lancaster said.

''With another intensive round of European games to come, including the Wasps-Harlequins and Sale-Clermont games on Sunday, there are likely to be additions to the squad after the weekend.

''There is plenty to be decided before the New Zealand match comes around.''

Manu Tuilagi has been omitted from the 33-strong party, but tellingly he has also been left out of the names not considered for squad selection due to injury.

Tuilagi was on Tuesday ruled out of all four of next month's Tests by his club Leicester because of a groin problem, but it is clear England hope he can yet play a role.

''We have included Jonathan Joseph as an injury replacement for Manu,'' Lancaster said.

''It's obviously disappointing to lose Manu but Jonathan has been in great form and deserves this chance.

''Our medics will continue to work closely with Leicester's and ensure that Manu is supported in his rehabilitation from his injury.''

England senior squad (33) for forthcoming autumn internationals:

Forwards: J Marler (Harlequins), M Mullan (Wasps), K Brookes (Newcastle), D Wilson (Bath), D Hartley (Northampton), R Webber (Bath), D Attwood (Bath), G Kruis (Saracens), J Launchbury (Wasps), C Lawes (Northampton), C Clark (Northampton), J Haskell (Wasps), B Morgan (Gloucester), C Robshaw (Harlequins, capt), B Vunipola (Saracens), T Wood (Northampton).

Backs: D Care (Harlequins), L Dickson (Northampton), B Youngs (Leicester), O Farrell (Saracens), G Ford (Bath), S Myler (Northampton), B Barritt (Saracens), L Burrell (Northampton), K Eastmond (Bath), J Joseph (Bath), B Twelvetrees (Gloucester), M Brown (Harlequins), A Goode (Saracens), J May (Gloucester), J Nowell (Exeter), S Rokoduguni (Bath), M Yarde (Harlequins).